Why are you acting like I said the word "negotiate"? I said "arrived at"; the means – friendly cooperation, terrain, a myriad individualist agreements, threat of brutal violence – are irrelevant to the overall point that the wolves have very obviously formed borders. Israel and Russia are no longer keeping the peace, hence they're actively dissolving the borders and redrawing them to whatever they can arrive at through military and geopolitical force.
I'll be one of the first people who'll tell you that animals experience real thoughts and emotions and have real, deep, complex bonds with their fellow animal. I'm sure these wolves' borders developed along natural formations, inherent population limits, intimidation, etc., and you'd have to be delusional to think these are totally divorced from the means by which states form borders.
You're treating this as an argument that borders are inherently good when the argument is that no true Scotsmanning the wolves is fucking bullshit. And if "keeping the peace" involves no (relatively) small-scale, cross-border violence, then I'd like to raise a point about wolves.